This paper reports on the genus Cobbionema Filipjev, 1922 in Sweden with the description of four species and a revision of the genus. Cobbionema acrocerca Filipjev, 1922 is relatively small in size, with a tail that has a conical proximal and a digitate distal section. Cobbionema cylindrolaimoides Schuurmans Stekhoven, 1950 is similar to C. acrocerca in most characters except having a larger body size and heavily cuticularized mandibles. Cobbionema brevispicula sp. nov. is characterised by short spicules and a conoid tail. Cobbionema acuminata sp. nov. is characterised by a long two-part spicule, a conical tail and three (one mid dorsal and two ventrosublateral) sharply pointed tines in the anterior chamber of the stoma that are located more anterior than in all the other species. We also present a molecular phylogeny of the family based on the nearly full-length 18S and the D2-D3 expansion segment of the 28S rRNA genes. Maximum Likelihood and Bayesian trees inferred from both genes strongly support a clade that included Cobbionema, Demonema Cobb, 1894 and Halichoanolaimus de Man, 1888 and another clade with Gammanema Cobb, 1920 and Latronema Wieser, 1954 nested together. None of the trees supported the monophyly of the subfamilies Choniolaiminae and Selachinematinae.
We review the species of Bidessus of Madagascar and describe Bidessus anjozorobe sp. nov. from material collected in Anjozorobe forest. Anjozorobe is part of the Anjozorobe-Angavo Protected Area, which is an important corridor of transition forest between typical eastern humid forests and the residual sub-humid forest of the Central Highlands. Bidessus longistriga Régimbart, 1895 and Bidessus perexiguus Kolbe, 1883 are widespread but endemic low-altitude species on Madagascar. Bidessus nesioticus Guignot, 1956 is an alpine species described from near the peak of the Ankaratra mountain massifs at 2500 m a.s.l. We recollected the species for the first time since its description, in Ankaratra and in a new area above 2000 m a.s.l. in the Andringitra mountain further south. Bidessus cf. nero Gschwendtner, 1933 is tentatively recorded for Madagascar for the first time but further studies are needed to test the status of mainland and insular populations. Bidessus apicidens Biström & Sanfilippo, 1986 has not been recollected on Madagascar since 1970. All species are endemic to Madagascar except potentially Bidessus cf. ceratus and Bidessus cf. nero described from Uganda and the Democratic Republic of the Congo, respectively. The older records of the two non-endemic species Bidessus complicatus Sharp, 1904 and Bidessus ovoideus Régimbart, 1895 on Madagascar could not be verified.
A new species of Paracrobeles, P. kelsodunensis sp. nov. is described from the Kelso Dunes area, Mojave National Preserve, southern California. Paracrobeles kelsodunensis sp. nov. is particularly characterised by a body length of 469–626 μm in females and 463–569 μm in males; lateral field with four incisures, extending almost to tail terminus; three pairs of asymmetrical lips, separated by U-shaped primary axils with two long guarding processes, each lip usually with four tines along its margin; three long labial probolae, deeply bifurcated, with slender prongs without tines; metastegostom with a strong anteriorly directed dorsal tooth; pharyngeal corpus anteriorly spindle-shaped, posteriorly elongate bulbous with dilated lumen; spermatheca 24–87 μm long; postvulval uterine sac 60–133 μm long; vulva in a sunken area; spicules 33–38 μm long; and male tail with a 5–8 μm long mucro. The generic diagnosis is emended on the basis of recently described species and a key to the species of Paracrobeles is provided.
Populations of Stegelleta are described from California, New Zealand and Senegal. An amphimictic population from California is identified as belonging to S. incisa and compared with type specimens from Utah and an amphimictic population from Italy. One population from New Zealand is close to S. incisa but considered to represent a new species, Stegelleta laterocornuta sp. nov. It is particularly characterised by a 379–512 μm long body in females and 365–476 μm in males; cuticle divided into 16 rows of blocks at midbody (excluding lateral field); lateral field with four incisures; three pairs of asymmetrical lips, U-shaped primary axils without guarding processes, each lip asymmetrically rectangular with a smooth margin, only lateral lips have slender acute tines; three labial probolae, bifurcated at half of their length; vulva without flap; spermatheca 17–31 μm long; postuterine sac 7–24 μm long; spicules 21.5–23.5 μm long. Other specimens from New Zealand are identified as belonging to S. tuarua. A parthenogenetic population from Senegal is identified as belonging to S. ophioglossa and compared with type specimens from Mongolia and records of several other populations of S. ophioglossa. The generic diagnosis is emended and a key to the species of Stegelleta is provided.
Oxyethira Eaton, 1873 is one of the most diverse genera of Hydroptilidae, comprising over 240 species distributed in all biogeographical regions. Here three new species of Oxyethira (Trichoglene) Neboiss, 1977 are described and illustrated from male specimens collected in New Caledonia: O. (Trichoglene) hamus sp. nov., recognized by the hook-shaped apex of the long inferior appendages in lateral view and by the posterior margin of segment IX with a trilobed appearance in ventral view; O. (Trichoglene) rectangulata sp. nov., recognized by the rectangular shape of the inferior appendages, which are totally fused and with two pairs of small setae on the inner face; and O. (Trichoglene) spiralis sp. nov., recognized by the strongly curvilinear shape of the subgenital process in dorsal
Three new species of Caledomina Johanson, 2011 are described and illustrated for the first time: C. paniensis sp. nov., C. dorsospina sp. nov. and C. kohensis sp. nov. A key to the males of the four known species in the genus is given, as well as a map indicating their geographical distribution. When examining a phylogenetic hypothesis comprising the four species, we see an increased complexity in younger species compared to older species.
Two new species of Oecetis from Maliau Basin, Malaysian Borneo, are described for the first time, O. mesospina sp. nov. and O. apelqvisti sp. nov. These two new species bring the total number of Oecetis found on the island of Borneo up to 16.
A total of nine families of Trichoptera were identified from material collected in Malaise andlight traps in the western part of Nyungwe National Park, southwestern Rwanda, late October 2018.Included in the material was an undescribed species of Pisuliidae which is described herein as Silvatareslaetae Ngirinshuti & Johanson sp. nov. The new species adds to the six Pisuliidae species previouslyrecorded for the East African region, five endemic to Tanzania and one to Uganda. This study portraysthe first results of an ongoing survey on the Trichoptera fauna of Rwanda.
Several taxonomic groups within Empidoidea Latreille, 1809 have been subject to unclear phylogenetic assignments along with multiple parallel hypotheses causing difficulties in classification and morphological identification. This study reviews the internal classification of the Ragadidae and includes a diagnosis and description of all included subfamilies and genera based on the results of an analysis of morphological characters using maximum parsimony. Illustration of important characters and a key to all genera in the family is given. The genus Hormopeza Zetterstedt, 1838 is found to be most closely related to Anthepiscopus Becker, 1891 and Iteaphila Zetterstedt, 1838, and the subfamily Iteaphilinae Wahlberg & Johanson, 2018 is therefore expanded to also include that genus. Hormopeza is consequently excluded from Ragadinae Sinclair, 2016. This study provides diagnoses, descriptions and keys in a contribution to a thorough classification of the empidoid groups and increased ease in morphological recognition.