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  • 1. Denk, Thomas
    et al.
    Tekleva, Maria V.
    Pollen morphology and ultrastructure of Quercus with focus on Group Ilex (=Quercus Subgenus Heterobalanus (Oerst.) Menitsky): implications for oak systematics and evolution2014In: Grana, ISSN 0017-3134, E-ISSN 1651-2049, Vol. 53, p. 255-282Article in journal (Refereed)
  • 2.
    Friis, Else Marie
    et al.
    Swedish Museum of Natural History, Department of Paleobiology.
    Mendes, Mário Miguel
    Universidade de Evora, Portugal.
    Pedersen, Kaj Raunsgaard
    University of Århus, Denmark.
    Paisia, an Early Cretaceous eudicot angiospermflower with pantoporate pollen from Portugal2018In: Grana, ISSN 0017-3134, E-ISSN 1651-2049, Vol. 57, p. 1-15Article in journal (Refereed)
    Abstract [en]

    A new fossil angiosperm, Paisia pantoporata, is described from the Early Cretaceous Catefica mesofossil flora, Portugal, based on coalified floral buds, flowers and isolated floral structures. The flowers are actinomorphic and structurally bisexual with a single whorl of five fleshy tepals, a single whorl of five stamens and a single whorl of five carpels. Tepals, stamens and carpels are opposite, arranged on the same radii and tepals are involute at the base clasping the stamens. Stamens have a massive filament that grades without a joint into the anther. The anthers are dithecate and tetrasporangiate with extensive connective tissue between the tiny pollen sacs. Pollen grains are pantoporate and spiny. The carpels are free, apparently plicate, with many ovules borne in two rows along the ventral margins. Paisia pantoporata is the oldest known flower with pantoporate pollen. Similar pantoporate pollen was also recognised in the associated dispersed palynoflora. Paisia is interpreted as a possibly insect pollinated, herbaceous plant with low pollen production and low dispersal potential of the pollen. The systematic position of Paisia is uncertain and Paisia pantoporata most likely belongs to an extinct lineage. Pantoporate pollen occurs scattered among all major groups of angiosperms and a close match to the fossils has not been identified. The pentamerous floral organisation together with structure of stamen, pollen and carpel suggests a phylogenetic position close to the early diverging eudicot lineages, probably in the Ranunculales.

  • 3.
    Friis, Else Marie
    et al.
    Swedish Museum of Natural History, Department of Paleobiology.
    Pedersen, Kaj Raunsgaard
    Crane, Peter Robert
    Welwitschioid diversity in the Early Cretaceous: Evidence from fossil seeds with pollen from Portugal and eastern North America2014In: Grana, ISSN 0017-3134, E-ISSN 1651-2049, Vol. 53, p. 175-196Article in journal (Refereed)
  • 4.
    Friis, Else Marie
    et al.
    Swedish Museum of Natural History, Department of Paleobiology.
    Pedersen, Kaj Raunsgaard
    Marone, Federica
    Arcellites punctatus sp. nov.: a new megaspore from the Early Cretaceous of Portugal studied using high resolution synchrotron radiation x-ray tomographic microscopy (SRXTM)2014In: Grana, ISSN 0017-3134, E-ISSN 1651-2049, Vol. 53, p. 91-102Article in journal (Refereed)
  • 5. Grímsson, F.
    Meller, Barbara (Contributor)
    Bouchal, Johannes M. (Contributor)
    Swedish Museum of Natural History, Department of Paleobiology.
    Grimm, Guido (Contributor)
    Combined LM and SEM study of the middle Miocene (Sarmatian) palynoflora from the Lavanttal Basin, Austria: Part III. Magnoliophyta 1 – Magnoliales to Fabales2015In: Grana, ISSN 0017-3134, E-ISSN 1651-2049, Vol. 54, p. 85-128Article in journal (Refereed)
    Abstract [en]

    Previous studies on the palynoflora from the Lavanttal Basin show that it contains a rich

    assemblage of spores and gymnosperm pollen. Present and ongoing investigations of

    dispersed angiosperm pollen suggest a high diversity within this group, and due to the

    excellent preservation of the material some rare pollen types are recognized. The Magnoliales

    to Fabales pollen record documented here contains 30 different taxa. Only a few pollen types

    are assigned to Magnoliids (four taxa); these are rare in the pollen record. Similarly, the

    Commelinids comprise five taxa and are also rare. Most of the angiosperm pollen originate

    from Eudicots, 21 taxa. Of the angiosperm taxa documented here, Magnolia , Carex ,

    Ranunculaceae, Platanus , Trochodendron , Buxus , Cercidiphyllum , Daphniphyllum ,

    Distylium , Fortunearia , Parrotia , Parthenocissus , Vitis , Euphorbia , Salix , and

    Papilionoideae are recorded for the first time from the Lavanttal Basin. This also includes the

    first fossil pollen record of Trochodendron  worldwide and the first reliable pollen record of

    Daphniphyllum . Several of the taxa described here had a wide Northern Hemispheric

    distribution from Eocene until the end of the Miocene. Also, key relatives of the fossil taxa

    are presently confined to humid warm-temperate environments, suggesting a very mild

    climate during the middle Miocene (Sarmatian) of the Lavanttal area. Some of the taxa

    encountered also support previous observations that the sediments of the Lavanttal Basin

    accumulated in a lowland wetland environment. This is based on pollen from aquatic taxa

    thriving in lakes, streams and swamps, and pollen of terrestrial plant taxa occupying margins

    of lakes and streams, backswamps, floodplains, river plains, and hummocks. Other 

     angiosperm pollen clearly originate from plants thriving on drier substrates, reflecting various

    vegetation units of the mixed evergreen/deciduous broad-leaved/conifer forests surrounding

    the wetland basin.

  • 6. Grímsson, F.
    Bouchal, Johannes M. (Contributor)
    Swedish Museum of Natural History, Department of Paleobiology.
    Meller, Barbara (Contributor)
    Zetter, Reinhard (Contributor)
    Combined LM and SEM study of the middle Miocene (Sarmatian) palynoflora from the Lavanttal Basin, Austria: part IV. Magnoliophyta 2 – Fagales to RosalesIn: Grana, ISSN 0017-3134, E-ISSN 1651-2049Article in journal (Refereed)
    Abstract [en]

    An ongoing investigation of the middle Miocene (Sarmatian) palynoflora from the Lavanttal Basin continues to show that it contains an extremely rich assemblage of angiosperm taxa. The Fagales to Rosales pollen record documented here contains 34 different taxa belonging to the Betulaceae (Alnus, Betula, Carpinus, Corylus, Ostrya), Fagaceae (Castanea, Fagus, Quercus Groups Cerris, Ilex, Cyclobalanopsis, Quercus/Lobatae), Juglandaceae (Engelhardioideae, Carya, Juglans, Pterocarya), Myricaceae (Morrella vel Myrica), Cannabaceae (Celtis), Elaeagnaceae (Elaeagnus), Rhamnaceae, Rosaceae (Prunus) and Ulmaceae (Cedrelospermum, Ulmus, Zelkova). Two of the pollen types represent extinct genera, Trigonobalanopsis and Cedrelospermum, and are also reported for the first time from the Lavanttal Basin along with pollen of Rhamnaceae and Prunus. The different types of Quercus pollen are now affiliated with Groups Cerris, Cyclobalanopsis, Ilex and Quercus/Lobatae based on sculpturing elements observed using scanning electron microscopy (SEM). Köppen signatures of potential modern analogues of the fossil Fagales and Rosales suggest a subtropical (Cfa, Cwa) climate at lower elevation and subsequent subtropical to temperate climate with altitudinal succession (Cfa→Cfb/Dfa→Dfb; Cwa→Cwb→Dwb) in the Lavanttal area during accumulation of the palynoflora. Most of the fossil taxa have potential modern analogues that can be grouped as nemoral and/or merido-nemoral vegetation elements, and the diversity of Fagales indicates a varying landscape with a high variety of niches.

  • 7. Mendes, Mário Miguel
    et al.
    Grimm, Guido
    Pais, João
    Friis, Else Marie
    Swedish Museum of Natural History, Department of Paleobiology.
    Fossil Kajanthus lusitanicus gen. et sp . nov. from Portugal: Floral evidence for Early Cretaceous Lardizabalaceae (Ranunculales, basal eudicot)2014In: Grana, ISSN 0017-3134, E-ISSN 1651-2049, Vol. 53, p. 283-301Article in journal (Refereed)
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